Mechanisms of control of locomotion and posture, visuomotor integration, synaptic plasticity & brain-machine interfacing.



2012 - 2015, Arizona State University, a student in the ASU-BNI  Interdisciplinary Graduate Program in Neuroscience

NSF graduate school fellow (CRFP)

                        Erik defended his Ph.D. on April 17th, 2015.


20082012    Arizona State University

College of Liberal Arts and Sciences: BS Mathematics, major; BS Economics, major;

School of Life Sciences: BS Biology, minor; BS Biochemistry, minor.


Honor thesis: Solving paralysis: a series of experiments on the contributions of pyramidal tract neurons to motor control.


2008-2010 Flinn Scholar

2011-2012 Goldwater Scholar


2004 - 2008,  Desert Vista High School, Phoenix, AZ. National Merit Scholar


Relevant Coursework:

Calculus I-II*, III; Advanced Calculus I

Differential Equations*; Linear Algebra

Statistics*; Probability; Applied Statistics; Mathematical Statistics

Stochastic Processes; Stochastic Modeling in Biology**

Mathematical Structures; Intermediate Real Analyses I

Object-oriented Program & Data

Physics: Mechanics, Electricity & Magnetism

Chemistry: Qualitative analysis*, Organic Chemistry I-II; Physical Chemistry

Biology I-II*, Organic evolution, Neurobiology, Cell Biology, Biochemistry, Genetics

Developmental Neurobiology**; Cellular and Molecular Neuroscience**; Systems Neuroscience**


* Completed in High School; ** Graduate Coursework


Research experience:

2007 – 2015     Research Assistant (paid, part-time), laboratory of Dr. Beloozerova

     Barrow Neurological Institute (BNI), Phoenix, AZ

2005 - 2007      High school student volunteer in the laboratory of Dr. Beloozerova, BNI


Science Outreach:

Sponsor/Mentor Desert Vista Science Club, 2008 –

Chaperone, Desert Vista HOSA Club, 2008 -

High School Student Mentor, BNI (SEPS) 2008 – 2015


Professional societies:

Member of: Society for Neuroscience




1.      Ertürk A, Mentz S, Stout EE, Hedehus M, Dominguez SL, Neumaier L, Krammer F, Llovera G, Srinivasan K, Hansen DV, Liesz A, Scearce-Levie KA, Sheng M. Interfering with the Chronic Immune Response Rescues Chronic Degeneration After Traumatic Brain Injury. J. Neurosci., 36(38): 9962-9975, 2016. PDF


2.      Stout EE, Sirota MG, Beloozerova IN. Known and unexpected constraints evoke different kinematic, muscle, and motor cortical neuron responses during locomotion. Eur. J. Neurosci., 42(9): 2666-2677, 2015. PDF


3.      Beloozerova IN, Stout EE, Sirota MG. Distinct thalamo-cortical controls for shoulder, elbow, and wrist during locomotion. Front. Comput. Neurosci., 7:62,  2013. Review. PDF


4.      Stout EE, Beloozerova IN. Differential responses of fast and slow conducting pyramidal tract neurons to changes in accuracy demands during locomotion. J. Physiol (L.), 591(Pt 10): 2647-2666, 2013. PDF


5.      Stout EE, Beloozerova IN. Pyramidal tract neurons receptive to different forelimb joints act differently during locomotion J. Neurophysiol., 107(7): 1890-1903, 2012. PDF


6.      Zelenin PV, Deliagina TG, Orlovsky GN, Karayannidou A, Stout EE, Sirota MG, Beloozerova IN. Activity of motor cortex neurons during backward locomotion. J. Neurophysiol., 105(6):2698-2714, 2011. PDF


7.      Karayannidou A, Beloozerova IN, Zelenin PV, Stout EE, Sirota MG, Orlovsky GN, Deliagina TG. Activity of pyramidal tract neurons in the cat during standing and walking on an inclined plane. J. Physiol. (L.), 587(Pt 15): 3795-3811, 2009. PDF


Poster presentations:


1.      Beloozerova IN, Marlinski V, Stout EE, Sirota MG. Distinct thalamo-cortical controls for shoulder, elbow, and wrist during locomotion. Annual Meeting of Neural Control of Movements Society, 2013.


2.      Stout EE, Sirota MG, Beloozerova IN. Changing steps on a short notice: insight from kinematics. Soc. Neurosci. Meeting, 274.16, 2012.


3.      Beloozerova IN, Marlinski V, Stout EE, Sirota MG. Thalamo-cortical network in locomotion.  Soc. Neurosci. Meeting, 78.16, 2012.


4.      Stout EE, Beloozerova IN. The role of slow-conducting pyramidal tract neurons in precision stepping. Soc. Neurosci. Meeting, 290.15, 2010.


5.      Zelenin PV, Deliagina TG, Orlovsky GN, Karayannidou A, Stout EE, Sirota MG, Beloozerova IN. Activity of corticospinal neurons in the cat during different forms of locomotion. 7th FENS Forum of European Neuroscience. 053.12, 2010.


6.      Stout EE, Beloozerova IN. Do Pyramidal Tract Neurons (PTNs) with different somato-sensory receptive fields have distinct firing properties during locomotion? Soc. Neurosci. Meeting, 565.16, 2009.


7.      B.J. Farrell, E.E. Stout, M.G. Sirota, I.N. Beloozerova, B.I. Prilutsky. Accurate target stepping in the cat: the full-body mechanics and activity of limb muscles. Soc. Neurosci. Meeting, 860.7, 2008.


8.      A. Karayannidou, I.N. Beloozerova, P.V. Zelenin, E.E. Stout, M.G. Sirota, G.N. Orlovsky, and T. G. Deliagina. Participation of pyramidal tract neurons in control of standing and walking on inclined surface. Soc. Neurosci. Meeting, 860.3, 2008. link


9.      E.E. Stout. Mechanisms of balance during walking. Arizona Science and Engineering fair, 2008.